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Maize

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Maize

Illustration depicting both male and female flowers of maize
Scientific classification
Kingdom: Plantae

(unranked): Commelinids

Order: Poales

Family: Poaceae

Genus: Zea

Species: Z. mays

Binomial name
Zea mays
L.

Maize (Zea mays L. ssp. mays, pronounced /ˈmeɪz/; also known in many English-speaking countries as corn), is a grass domesticated by indigenous peoples in Mesoamerica in prehistoric times. The Aztecs and Mayans cultivated it in numerous varieties throughout central and southern Mexico, to cook or grind in a process called nixtamalization. Later the crop spread through much of the Americas. Between 1250 A.D. and 1700 A.D. nearly the whole continent had gained access to the crop. Any significant or dense populations in the region developed a great trade network based on surplus and varieties of maize crops. After European contact with the Americas in the late 15th and early 16th centuries, explorers and traders carried maize back to Europe and introduced it to other countries through trade. Its ability to grow in distinct climates, and its use were highly valued, thus spreading to the rest of the world.

Maize is the most widely grown crop in the Americas (332 million metric tons annually in the United States alone). Hybrid maize, because of its high-grain yield as a result of heterosis ('hybrid vigor')[citation needed], is preferred by farmers over conventional varieties[citation needed]. While some maize varieties grow up to 7 metres (23 ft) tall,[1] most commercially grown maize has been bred for a standardized height of 2.5 metres (8.2 ft). Sweet corn is usually shorter than field-corn varieties.

Naming conventions

Many small male flowers make up the male inflorescence, called the tassel

The term maize derives from the Spanish form of the indigenous Taino word maiz for the plant. This was the term used in the United Kingdom and Ireland, where it is now usually called "sweet corn", the most common form of the plant known to people there.[2]

Outside the British Isles, another common term for maize is "corn". This was originally the English term for any cereal crop. In North America, its meaning has been restricted since the 19th century to maize, as it was shortened from "Indian corn."[3] The term Indian corn now refers specifically to multi-colored "field corn" (flint corn) cultivars.[4]

In scientific and formal usage, "maize" is normally used in a global context. Equally, in bulk-trading contexts, "corn" is used most frequently. In the UK, Australia and other English-speaking countries, the word "corn" is often used in culinary contexts, particularly in naming products such as popcorn and corn flakes. "Maize" is used in agricultural and scientific references.[5]

In Southern Africa, maize is commonly referred to as mielie or mealie, from the Portuguese milho.[6] Mielie-meal is the ground form.

Structure and physiology

Female inflorescence, with young silk

Maize stems superficially resemble bamboo canes and the internodes can reach 20–30 centimetres (8–12 in). Maize has a distinct growth form; the lower leaves being like broad flags, 50–100 centimetres long and 5–10 centimetres wide (2–4 ft by 2–4 in); the stems are erect, conventionally 2–3 metres (7–10 ft) in height, with many nodes, casting off flag-leaves at every node. Under these leaves and close to the stem grow the ears. They grow about 3 millimetres a day.

The ears are female inflorescences, tightly covered over by several layers of leaves, and so closed-in by them to the stem that they do not show themselves easily until the emergence of the pale yellow silks from the leaf whorl at the end of the ear. The silks are elongated stigmas that look like tufts of hair, at first green, and later red or yellow. Plantings for silage are even denser, and achieve a lower percentage of ears and more plant matter. Certain varieties of maize have been bred to produce many additional developed ears. These are the source of the "baby corn" used as a vegetable in Asian cuisine.

Maize is a facultative long-night plant and flowers in a certain number of growing degree days > 50 °F (10 °C) in the environment to which it is adapted.[7] The magnitude of the influence that long nights have on the number of days that must pass before maize flowers is genetically prescribed and regulated by the phytochrome system.[8] Photoperiodicity can be eccentric in tropical cultivars, while the long days characteristic of higher latitudes allow the plants to grow so tall that they do not have enough time to produce seed before being killed by frost. These attributes, however, may prove useful in using tropical maize for biofuels.[9]

Stalks, ears, and silk
male flowers

The apex of the stem ends in the tassel, an inflorescence of male flowers. When the tassel is mature and conditions are suitably warm and dry, anthers on the tassel dehisce and release pollen. Maize pollen is anemophilous (dispersed by wind) and because of its large settling velocity most pollen falls within a few meters of the tassel. Each silk may become pollinated to produce one kernel of maize. Young ears can be consumed raw, with the cob and silk, but as the plant matures (usually during the summer months) the cob becomes tougher and the silk dries to inedibility. By the end of the growing season, the kernels dry out and become difficult to chew without cooking them tender first in boiling water. Modern farming techniques in developed countries usually rely on dense planting, which produces one large ear per stalk.

Seeds

The kernel of maize has a pericarp of the fruit fused with the seed coat, typical of the grasses, and the entire kernel is often referred to as the seed. The cob is close to a multiple fruit in structure, except that the individual fruits (the kernels) never fuse into a single mass. The grains are about the size of peas, and adhere in regular rows round a white pithy substance, which forms the ear. An ear contains from 200 to 400 kernels, and is from 10–25 centimetres (4–10 inches) in length. They are of various colors: blackish, bluish-gray, purple, green, red, white and yellow. When ground into flour, maize yields more flour, with much less bran, than wheat does. However, it lacks the protein gluten of wheat and, therefore, makes baked goods with poor rising capability and coherence.

A genetic variant that accumulates more sugar and less starch in the ear is consumed as a vegetable and is called sweet corn.[citation needed]

Immature maize shoots accumulate a powerful antibiotic substance, DIMBOA (2,4-dihydroxy-7-methoxy-1,4-benzoxazin-3-one). DIMBOA is a member of a group of hydroxamic acids (also known as benzoxazinoids) that serve as a natural defense against a wide range of pests including insects, pathogenic fungi and bacteria. DIMBOA is also found in related grasses, particularly wheat. A maize mutant (bx) lacking DIMBOA is highly susceptible to be attacked by aphids and fungi. DIMBOA is also responsible for the relative resistance of immature maize to the European corn borer (family Crambidae). As maize matures, DIMBOA levels and resistance to the corn borer decline.

Because of its shallow roots, maize is susceptible to droughts, intolerant of nutrient-deficient soils, and prone to be uprooted by severe winds.[10]

Some hazards of maize in the diet

Pellagra

When maize was first introduced into other farming systems than those used by traditional native-American peoples, it was generally welcomed with enthusiasm for its productivity. However, a widespread problem of malnutrition soon arose wherever maize was introduced as a staple food. This was a mystery since these types of malnutrition were not normally seen among the indigenous Americans, to whom maize was the principal staple food.[11]

It was eventually discovered that the indigenous Americans learned long ago to soak maize in alkali-water—made with ashes by North Americans and lime (calcium oxide) by Mesoamericans—which liberates the B-vitamin niacin, the lack of which was the underlying cause of the condition known as pellagra. This alkali process is known by its Nahuatl (Aztec)-derived name: nixtamalization.

Besides the lack of niacin, pellagra was also characterized by protein deficiency, a result of the inherent lack of two key amino acids in pre-modern maize, lysine and tryptophan. Nixtamalisation was also found to increase the availability of lysine and tryptophan to some extent, but more importantly, the indigenous Americans had learned long ago to balance their consumption of maize with beans and other protein sources such as amaranth and chia, as well as meat and fish, in order to acquire the complete range of amino acids for normal protein synthesis.

Since maize had been introduced into the diet of non-indigenous Americans without the necessary cultural knowledge acquired over thousands of years in the Americas, the reliance on maize in other cultures was often tragic. In the late 19th century pellagra reached epidemic proportions in parts of the deep southern U.S., as medical researchers debated two theories for its origin: the deficiency theory (eventually shown to be true) posited that pellagra was due to a deficiency of some nutrient, and the germ theory posited that pellagra was caused by a germ transmitted by stable flies. In 1914 the U.S. government officially endorsed the germ theory of pellagra, but rescinded this endorsement several years later as evidence grew against it. By the mid-1920s the deficiency theory of pellagra was becoming scientific consensus, and the theory was proved in 1932 when niacin deficiency was determined to be the cause of the illness.